A locally common pioneer terricolous lichen on bare acidic soil (sandy soil, peat, humus), mainly on open stands (heathlands, raised bogs, screes, alpine habitats, forest openings, etc.), where it forms communities with its relatives (P. icmalea, P. uliginosa and Trapeliopsis granulosa). Rarely, it also colonizes wood. Within the genus, it is easily recognizable due to relatively large thallus granules or areoles (more than 0.1 mm). Juvenile specimens with small granules, potentially confusable with P. icmalea, are easily recognizable based on secondary metabolites (C- reaction). The species was not distinguished in the past, so only a revision of herbarium material under the epithets uliginosa, humosa or fuliginea would clarify its historical distribution in the Czech Republic.taxonomic classification:
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