An inconspicuous lichen similar to T. aromatica, distinguished by darker squamules without pale spots and by brown epihymenium. It mainly grows on calcareous rocks at higher elevations, usually on cyanolichen thalli, especially on species of Placynthium (Timdal 1992). The lichen is scattered all over Europe. There is a single published record from the Czech Republic – from the Suchý žleb valley in Moravský kras (Moravian Karst), with very atypical ecology; it was recorded growing on the bryophyte Schistidium apocarpum (Servít 1910, as T. aromatica var. cervina) and thus, requires revision. A subrecent unpublished collection comes from crystalline limestones in Rudník in Obří důl in the Krkonoše Mts (Palice, unpubl.).
Literature: Timdal E. (1992): A monograph of the genus Toninia (Lecideaceae, Ascomycetes). – Opera Botanica 110: 1–137. Servít M. (1910): První příspěvek k lichenologii Moravy. – Zprávy Kommisse pro Přírodovědecké Prozkoumání Moravy, oddělení botanické, Brno, 6: 1−83.taxonomic classification: most frequented synonyms:Toninia verrrucariodes
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